The best replacement for underlining is quotation marks.

The best alternatives to underlining are rated as following:
★★★★★  quotation marks
★★★★  italics
★★  quotations
★★  quotes
★★  underlying
★★  italicizing
★★  highlighting
★★  lining
★  quoting
  interlining
  quotation

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No companions for underlining have been suggested so far.
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Google Trends

Scale is based on the average worldwide traffic of underlining in all years. Click on graph for more.
underlining
quotation marks
italics