The best replacement for learned is learnt.

Лучшие альтернативы к learned можно упорядочить следующим образом:
★★★★★  learnt
★★★  inherited traits
★★  innate behavior
★★  inherited
★★  innate
★★  learnt grammar
★  inherited behaviors
★  instinctive behavior
★  educated
★  learned
★  have learned
★  taught
★  instinctive
★  learn-ed
  inherited behavior
  have learnt
  learnt english
  learnt uk
  learnt define
  instinct
  acquired
  unlearned
  are leaders born
  i learned
  innate behaviour
  lessons learnt
  instinctive behaviors
  lesson learnt
  leant
  innate behaviors
  learner
  instinct behavior

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