The best replacement for species 8472 is dominion.

The best alternatives to species 8472 are rated as following:
★★★★★  dominion
★★★★  vorlons
★★★★  q
★★★  borg
★★★  yuuzhan vong
★★★  shadows
★★  hirogen
★★  voth

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No companions for species 8472 have been suggested so far.
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species 8472
dominion
vorlons